Sara sacrifices herself for the safety of Michael so the brothers can get on with their final plan to attain freedom, but their plan gets ‘shipwrecked’. Meanwhile, Bellick blackmails Sucre to partner up as T-Bag’s streak of murder continues to the south of the border.
Westmoreland wants Michael to speed up the planned escape, and gets Bellick one step closer to unraveling it. Michael has to decide whether to involve Sara in the breakout plan. Tweener’s devotion is put to a test. Veronica finds out that her ally unexpectedly has a connection to an inmate serving time at Fox River.
Pandemonium erupts at the hotel following the assassination, and Christina puts the final pieces of her plan together. Meanwhile, the General makes good on his threat, and Michael is forced to choose between saving either Lincoln or Sara.
Hades turns to the heroes to ask for help with getting Zelena back from Rumplestiltskin and Peter Pan. In return, he offers to take all their names off their tombstones. However, when Hook is still unable to leave, he and Emma must journey into the depths of the Underworld. Meanwhile, Cruella De Vil is determined to keep the heroes trapped in the Underworld. In flashbacks, Emma searches for answers about her family and makes an unexpected friendship.
With Mary Margaret in labor and due to give birth at any moment, the residents of Storybrooke go on high alert in an attempt to ensure that Zelena doesn’t try to steal her newborn baby for use in her ultimate game plan to turn back time and change her destiny — while obliterating Regina’s existence. Meanwhile, in the land of Oz of the past, Glinda tries to convince Zelena to fight her evil tendencies and join her and her sister witches as a protector of Oz. But the appearance of a young girl from Kansas could turn out to be her undoing.
Emma’s potential for darkness is looming over everyone, but when Emma realizes Maleficent’s daughter Lily is in fact her closest friend from her foster care days, she resolves to find Lily and reunite her with her mother. Regina joins forces with Emma and together they set out to track down Lily and to warn Robin about Zelena. However, neither of them is prepared for the harsh realities they’ll encounter in the outside world. Meanwhile, in Storybrooke, Gold faces a crisis involving Belle. In a foster care flashback, things are looking up for young Emma with her new family until Lily’s appearance threatens to destabilize everything.
Hook helps Regina with a plan to transport her and Henry back to Fairytale land; Emma grows increasingly suspicious of Tamara; the queen asks Rumpelstiltskin to transform her so that she can kill Snow White.
August tells Emma how to beat Regina and how she can get custody of Henry, while Regina hatches a plot to seduce David now that Mary Margaret is back at work. In the fairy-tale world, a plan to save Snow White and Prince Charming’s daughter unfolds and is agreed to by Geppetto, who has his own child’s safety in mind.