Rebecca’s relationships are tested as she tries to navigate her friendship with Paula, her history with Josh and her connection with Greg. While at Josh’s sister Jayma’s wedding, Rebecca’s world is turned upside down.
Liv and Ravi are in need of help and find an unlikely ally in Blaine. Detective Babineaux and Dale Bozzio pursue a new lead on an old case. Meanwhile, when Major finds himself in a world of trouble, sleazebag Brant Stone comes to his rescue. Vaughn’s evil side reaches a whole new level of mean.
In an effort to recreate the missing piece of the tattoo, Michael gets transferred to the psych ward, seeking the help of Haywire. T-Bag and C-Note have to join forces.
Michael and Lincoln come to blows over Scylla as Christina sets the wheels in motion for her plan. Meanwhile, Sara receives life-changing news. T-Bag arrives at the Indian Embassy. The General becomes increasingly paranoid.
T-Bag digs deep and finds his ‘nice-side’ with the help of a psychiatrist. Michael, Lincoln, and Sarah get their hands on evidence needed to bring down The Company, while Kellerman is close to finding the President.
In flashbacks, Ruby and Mulan find themselves in Oz, where they meet Dorothy. After the three witness Zelena’s return to Oz, they look for a way to defeat her once and for all. However, Dorothy mysteriously disappears, and Ruby’s search for her new friend lands her in the Underworld. Reunited with the heroes, Ruby teams up with Emma, Regina and Snow to continue looking for Dorothy. Meanwhile, Snow and David struggle with not being able to be with their son, Neal, and devise a plan so that one of them can escape the Underworld.
While off on her own to deal with what happened with Cora, Mary Margaret comes across an entirely wooden August; Neal invites his fiancee to Storybrooke.
Emma is still reeling after learning the truth of her parents’ history with Maleficent, but she must focus on finding the Author, who is running loose in Storybrooke, before Gold does. When Gold’s quest for his happy ending grows more urgent, he blackmails Regina into helping him. In a Fairy Tale Land flashback, Robin Hood learns what it means to be an honorable thief when he accepts a proposition by Rumpelstiltskin to travel to Oz to steal a valuable magic elixir from the Wicked Witch.
After Zelena steals Regina’s heart, Regina casts a spell so that she can speak across the realms to her dead mother, Cora, to discover the truth about why she abandoned Zelena, and Belle stumbles across what Zelena’s ultimate end game is. Meanwhile, in the Fairy Tale Land that was, young Cora is duped by a man claiming to be a prince and finds herself alone and pregnant. But a chance meeting with a real prince could lead Cora to the royal life she’s always craved, but she must keep her pregnancy a secret or risk losing everything.
Emma continues her exhaustive search for evidence that will prove Mary Margaret’s innocence in the murder of David’s wife, Kathryn. Meanwhile, in the fairytale land that was and before evil blackened her soul, Regina must choose between betraying her mother, Cora, and marrying for true love, or betrothing royalty and living a regal – but loveless – life; and the event that caused the Evil Queen to loathe Snow White is revealed.